Ammonia from their urine can actually kill them....
not sure how often you clean their cages, but I'd recommend at the very least once a week. Personally I clean mine out twice, somtimes 3 times a week, just to make sure they're all comfy. Ammonia is very deadly, even if only inhaled in small amounts, remember the smell of urine is actually harmful because of ammonia, it gives some people bad allergic reactions etc. but these are little bitty animals and it only takes a fraction of a given substance to irritate them. In essence what i'm saying is....if it (smell, look of the cage, lack of toys) seems bad to you, it's about 10 times worse if you were the rats/mice/gerbils etc 'cause they have to live in it. Other than ammonia, i'm not sure what could have caused it linds, you've already ruled out the mother nursing and they'd been getting milk. Best of luck.
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